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Libellula composita tandem pair.  Photo by Roy Beckemeyer, New Mexico, July 2001.Tandem Flight and Oviposition Flight in Libellula composita (Odonata: Anisoptera: Libellulidae)   Photo above right: Libellula composita pair in tandem flight

A Web Page by Roy J. Beckemeyer     

Last Updated 29 January 2004


While in the BLM Overflow Area adjacent to Bottomless Lakes State Park in Chaves County, New Mexico in July of 2001, I had the opportunity to observe males, females, and tandem pairs of The Bleached Skimmer (Libellula composita) in flight.  I used a 200 mm Nikon macro lens and a flash to take a series of pictures of the species in flight.  Although many of the images are not exceptionally clear, they are good enough to show some interesting facets of tandem flight and of method of oviposition in this species.  Some of the tandem flight pictures are accessible from my Natural Flight web page as well, but I wanted to group them together here so they could be compared more readily.  By clicking on any picture, it will be brought up to full screen (760 pixels wide) size.

Tandem flight of Libellula composita.  Photo by Roy Beckemeyer, July 2001, New Mexico.Tandem flight of Libellula composita.  Photo by Roy Beckemeyer, July 2001, New Mexico.

 

 

 

 

 

 

 

Tandem flight of Libellula composita.  Photo by Roy Beckemeyer, July 2001, New Mexico.In these three photos it is interesting to see how the female uses her legs to grip the male's abdomen.  The abdomen of the male is thus sort of wrapped around the head of the female.  Compare this to the pictures of Sympetrum corruptum, where the female does not grasp the male abdomen.  This is the formation in which the pair fly about from place to place between actual bouts of oviposition.

The male and female do not generally seem to be flapping with fore and hind wings of each in phase with those of the other, but the respective wing pairs are not very far out of phase either.


Pair of Libellula composita ovipositing in tandem.  Photo by Roy Beckemeyer, July 2001, New Mexico. Pair of Libellula composita ovipositing in tandem.  Photo by Roy Beckemeyer, July 2001, New Mexico.


In this pair of photos the male and female are both hovering, with their bodies held nearly vertically, so that the female can lay her eggs.  She does so by flicking the tip of her abdomen in the water to wash off the small mass of eggs that has accumulated on the underside of her terminal abdominal segments.  In the picture at right above, the water droplets that she has flipped up in the process of ovipositing can be seen near the bottom center of the picture, showing how forcefully she has flicked her abdomen.

Females in this species also oviposit alone at times.  The next two photos show a single female ovipositing.  One is a front view, the other a side view.  In both pictures, the water scooped up off the surface and propelled by the flick of her abdomen can be seen.  Note that in the picture to the left, the jet of water is wavy or sinusoidal - this indicates that, as the female's abdomen tip was traversing the water, she was oscillating in a rolling motion.  This is a neat indication of the complicated dynamics of dragonfly flight.  I have written a paper on this that will appear in Odonatologica in 2004.

 

Female Libellula composita laying eggs and flipping water droplets in the process.  Photo by Roy Beckemeyer, July 2001, New Mexico.Female Libellula composita laying eggs and flipping water droplets in the process.  Photo by Roy Beckemeyer, July 2001, New Mexico.


Scanned image of ventral view of female Libellula composita showing vulvar lamina and flanged area of 8th segment.At right is a scanned image of the underside of a female Libellula composita.  It shows the shape and appearance of the ventral side of the terminal abdominal segments, including the vulvar lamina and the flanged area of the 8th abdominal segment.  It is provided to aid in interpreting the oviposition photos, in two of which this area of the abdomen can be clearly seen.

 


Click HERE to link to a similar page on Sympetrum corruptum

Click HERE to link to Takashi Aoki's Page of Movies of Odonata Ovipositing


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