CHECKLIST OF Midco, Oklahoma PERMIAN INSECTS
Orders in taxonomic sequence per
Carpenter (1992):
"Treatise
on Invertebrate Paleontology. Part R. Arthropoda 4. Vol. 3 & 4.
Hexapoda,"
Taxonomic assignments and synonymies per that reference
as well.
Synonyms given when they relate to names used in the works of
Sellards, Tillyard, Carpenter, and Tasch & Zimmerman.
For other synonyms,
see the Treatise volumes.
A Web Page by Roy J. Beckemeyer
Revised 14/1/11: Added new genus Liomopterella, (Beckemeyer, R. J., 2010, A new species of Liomopterella Sharov, 1961 (Polyneoptera: Liomopteridae) from the Early Permian Wellington Formation of Noble County, Oklahoma, Trans. Kansas Acad. Sci. 113(3-4): 151-160), revised Homocladus Bethoux, Beckemeyer, Engel and Hall, 2010, New data on Homocladus grandis, a Permian stem-mantodean (Polyneoptera: Dictyoptera), Journal of Paleontology, 84(4): 746-753), revised familial assignment of Elmopterum (Aristov, D. S., 2009, Review of the stratigraphic distribution of Permian grylloblattida (Insecta), with descriptions of new taxa, Paleontological Journal, 43(6): 643-651).
14 September 2009. Corrected error in familial assignment of genera Parelmoa and Pseudelmoa (Diaphanopterodea: Parelmoidae) pointed out to me by David Nicholson. Added Diaphnopterodea: Biarmohymenidae: Anomalohymen dochmus Beckemeyer and Engel, 2009. Removed Diaphanopterodea: Elmoidae: Elmoa trisecta from this list as it has been reported only for Elmo.
8 August, 2007 - Corrected delinquency by adding in Coleoptera entry for Lubkin & Engel, 2005.
22 July 2007 - Added taxa from Béthoux & Beckemeyer 2007, identified 2003 Orthoptera species.
15 Nov. 2003 - added new specimen: New family: new genus: new species of Orthoptera.
15 April 2004 - added names of 2 new taxa: Lophioneuridae: Cyphoneurodes patriciae Beckemeyer, 2004 and Diaphanopterodea: Martynoviidae: Martynovia halli Beckemeyer, 2004.
2 May, 2004 - added names of 3 new taxa: Blattinopsidae: Glaphyrophlbia ovata (Sellards) 1909, Stereopteridae: Stereopterum rotundum Carpenter, 1950, and Mecoptera: Permopanorpidae: Permopanorpa inaequalis Tillyard, 1926.
7 Aug. 2004 - added names of Raaschiidae: Raaschia oklahomensis Beckemeyer, 2004, Caloneurodea: Gigagramma carpenteri Bethoux, Nel & Lapeurie, 2004, Caloneurodea: Paleuthygramma acuata Carpenter, 1943 [per Bethoux, Nel & Lapeyrie, 2004], Caloneurodea: Anomalogramma parva Carpenter, 1943 [per Bethoux, Nel & Lapeyrie, 2004], Uncertain Order (Caloneurodea or Orthoptera): Lipogramma sinuosa Bethoux, Nel & Lapeyrie, 2004.
Use your web browser "Find" function to search the page for known taxa.
Key:
No indentation:
Order (Higher Taxon) Geological time of occurrence (Number of genera,
species)
One tab: Suborder
Author Year described [synonymies in brackets]
Two tabs: Family Author Year
described [synonymies in brackets]
Three tabs: Genus Author Year
described [synonymies in brackets]
Four tabs: species Author (in parentheses
if described in another Genus) Year described (Original
Genus)
[synonymies in brackets]
Archaeognatha (Apterygota) U. Carb. - Hol. (1 family, 1 genus, 1 species)
Uncertain family
Uncertain genus
Uncertain species (specimens observed in Oklahoma State University collection by Roy Beckemeyer, 2010)
Ephemeroptera (Pterygota: Palaeoptera) U.Carb.-Hol. (2 families, 2 genera, 7 species) [=Plectoptera]
Misthodotidae Tillyard 1932
Misthodotes Sellards 1909 [nom. subst. pro Dromeus Sellards 1907;= Eudoter Tillyard 1936]
obtusus (Sellards) 1909 (Dromeus)
edmundsi Carpenter 1979
ovalis Tillyard 1932
Protereismatidae Lameere 1917 [= Protereismopheridae Sellards 1907]
Protereisma Sellards 1907 [= Protechma, Prodromus, Bantiska, Pinctodia, Rekter Sellards, 1907; Esca, Mecus Sellards 1909; Loxophlebia Martynov 1928]
arcuatum (Sellards) 1907 (Rekter)
latum Sellards 1907
directum Carpenter 1979
americanum (Demoulin) 1970 (Kukalova)
Palaeodictyoptera (Pterygota: Palaeoptera) U.Carb.-Perm. (1 family, 1 genus, 1 species)
Calvertiellidae Martynov 1931
Moravia Kukalova 1964
grandis Carpenter 1979
Megasecoptera (Pterygota: Palaeoptera) U.Carb.-Perm. (1 family1 genus, 9 species) [= Protohymenoptera Tillyard 1924]
Protohymenidae Tillyard 1924
Protohymen Tillyard 1924 [= Pseudohymen Martynov 1932, Pseudohymenopsis Zalessky 1956]
readi Carpenter 1933
curvatus Carpenter 1947
latus Carpenter 1947
venustus Carpenter 1947
largus Carpenter 1947
bifurcatus Zimmerman in Tasch and Zimmerman 1962
shafferi Zimmerman in Tasch and Zimmerman 1962
pictus Zimmerman in Tasch and Zimmerman 1962
anomalus Zimmerman in Tasch and Zimmerman 1962
Diaphanopterodea (Pterygota: Palaeoptera) U.Carb.-Perm. (5 families, 9 genera, 19 species)
Asthenohymenidae Tillyard 1924
Asthenohymen Tillyard 1924 [= Karoohymen Riek 1976]
dunbari Tillyard 1924 [= A. gracilis, A. kansasensis, A. stenobasis, Tillyard, 1926; Doter minor Carpenter 1930]
pusillus Tillyard 1926 [= Doter pusillus Carpenter 1930]
triangularis Carpenter 1947
apicalis Carpenter 1947
parvulus Carpenter 1947
latus Carpenter 1947
minutus Zimmerman in Tasch and Zimmerman 1962
Parelmoidae Rohdendorf 1962
Parelmoa Carpenter 1947
revelata Carpenter 1947
radialis Carpenter 1947
obtusa Carpenter 1947
Pseudelmoa Carpenter 1947
ampla Carpenter 1947
Bardohymenidae Zalessky 1937
Calohymen Carpenter 1947
permianus Carpenter 1947
Sylvohymen Carpenter 1947
ingens Carpenter 1947
Biarmohymenidae Zalessky 1937
Anomalohymen Beckemeyer and Engel 2009
dochmus Beckemeyer and Engel 2009
Martynoviidae Tillyard 1932 [= Martynoviella Tillyard 1932]
Martynovia Tillyard 1932
longipennis Carpenter 1947
halli Beckemeyer 2004
Eumartynovia Carpenter 1947
raaschii Carpenter 1947
Phaneroneura Carpenter 1947
martynovae Carpenter 1947
reducta Carpenter 1947
Protodonata (Pterygota: Palaeoptera) U.Carb.-Perm. (2 families, 3 genera, 3 species) [= Meganisoptera Martynov 1932]
Meganeuridae Handlirsch 1906
Meganeuropsis Carpenter 1939
ameriana Carpenter 1947
Tupus Sellards 1906 [= Typus Sellards 1909, unjustified emendation; Meganeurela Handlirsch 1906 among others]
gracilis Carpenter 1947
Paralogidae Handlirsch 1906
Oligotypus Carpenter 1931
Odonata (Pterygota: Palaeoptera) Perm.-Hol. (2 families, 3 genera, 6 species)
Archisozygoptera Handlirsch 1906 [= Protozygoptera Tillyard 1925]
Kennedyidae Tillyard 1925
Kennedya Tillyard 1925
mirabilis Tillyard 1925
fraseri Carpenter 1947
Progoneura Carpenter 1931
nobilis Carpenter 1947
venula Carpenter 1947
sp. Carpenter 1947
Protanisoptera Carpenter 1931
Ditaxineuridae Tillyard 1926
Ditaxineura Tillyard 1926
celulosa Carpenter 1933
Protorthoptera (Pterygota: Neoptera: Exopterygota) U.Carb.-Triass. (9 families, 13 genera, 14 species) [= Protoperlaria Tillyard 1928; Protoblattoidea Handlirsch 1906, Paraplecoptera Martynov 1938, among others]
Blattinopsidae Bolton 1925 [= Oryctoblattina Scudder 1879 among others]
Glaphyrophlebia Handlirsch 1906 [= Pursa, Sindon, Sellards, 1909]
speciosa (Sellards) 1909 (Sindon) - in the collection of Roy Beckemeyer - a specimen that had been collected by Tasch in 1959 but not previously reported, and a specimen collected by Joseph Hall from Raasch 5 in 2003
ovata (Sellards) 1909 (Pursa) - in the collection of Roy Beckemeyer - a specimen collected by Joseph Hall from Raasch 5 in 2004
Chelopteridae Carpenter 1950
Chelopterum Carpenter 1950
peregrinum Carpenter 1950 [not listed by Carpenter or by Tasch & Zimmerman; found in ex Tasch collection material by Roy J. Beckemeyer in March, 2000 - see Midco Home page]
Lemmatophoridae Sellards 1909 [= Ortaidae Sellards 1909, Lecoriidae handlirsch 1919]
Lemmatophora Sellards 1909
typa Sellards 1909 [= typica Tillyard 1928; anomala, elongata, delicosa, hirsuta, Sellards 1909]
Artinska Sellards 1909 [= Estadia, Lectrum, Orta, Sellards 1909]
clara Sellards 1909 [= medialis, gracilis, pecta, major, extensa, Sellards, 1909; Estadia elongata, arcuata, tenuis, Sellards, 1909; Estadia tripunctata Carpenter 1926; Lectrum anomalum Sellards, 1909]
Lisca Sellards 1909
minuta Sellards 1909 [= Lemmatophora reducta, minuta, Tillyard 1928; Lemmatophora cubitalis Handlirsch 1937]
Liomopteridae Sellards 1909
Liomopterella Sharov, 1961
pediaecetae Beckemeyer, 2010
Liomopterum Sellards 1909 [= Horates Sellards 1909]
ornatum Sellards 1909 [= L. extensum Sellards 1909]
Midcopterum Béthoux & Beckemeyer 2007
evidens Béthoux & Beckemeyer 2007
Phenopteridae Carpenter 1950
Phenopterum Carpenter 1950 [nom. subst. pro. Lepium Sellards 1909]
elongatum (Sellards) 1909 (Lepium) [= Lepium reticulatum Sellards 1909; Lepium sellardsi Handlirsch 1919]
Probnidae Sellards 1909 [nom. correct. pro Probnisidae Sellards 1909]
Probnis Sellards 1909 [= Espira, Stoichus, Stinus, Sellards 1909]
speciosa Tillyard 1937 [= P. coriacea, Espira obscura, Stoichus elegans, S. arcuatus, S. minor, S. tenuis, Stinus brevi-cubitalis, Sellards 1909; Probnis sellardsi Handlirsch 1919]
Raaschiidae Beckemeyer 2004
Raaschia Beckemeyer 2004
oklahomensis Beckemeyer 2004 [specimen found at TaschIIA, Noble Co., OK, May 2003 by Roy Beckemeyer & Joseph Hall]
Stereopteridae Carpenter 1950
Stereopterum Carpenter 1950
rotundum Carpenter 1950 - a specimen collected by Roy Beckemeyer from Billings 3 locality in 2004
Tshekardominidae Novokshonov and Aristov, 2002
Sigmophlebia Béthoux & Beckemeyer 2007
engeli Béthoux & Beckemeyer 2007
Blattaria (Pterygota: Neoptera: Exopterygota) U.Carb.-Hol. (1 family, 2 genera, 2 species)
Spiloblattinidae Handlirsch
Permoblattina Tillyard 1937
curta (Sellards) 1908 (Etoblattina)
Pycnoblattina Sellards 1908
curvata (Sellards) 1908 (Puknoblattina)
Protelytroptera (Pterygota: Neoptera: Exopterygota) Perm.-Cret. (1 family, 2 genera, 2 species)
Protelytridae Tillyard 1931
Protelytron Tillyard 1931
sp. Zimmerman in Tasch and Zimmerman 1962 (p. 1327)
Uncertain Family
Uncertain genus
sp. Zimmerman in Tasch and ZImmerman 1962 (p. 1327)
Orthoptera (Pterygota: Neoptera: Exopterygota) U.Carb.-Hol. (1 family, 2 genera, 2 species)
Oedischiidae Handlirsch 1906
Mezenoedischiinae Gorochov 1987
Pseudoiasvia Béthoux & Beckemeyer 2007
sinuosa Béthoux & Beckemeyer 2007
Undetermined Family
Crinoedischia Béthoux & Beckemeyer 2007
noblensis Béthoux & Beckemeyer 2007
Psocoptera (Pterygota: Neoptera: Exopterygota) Perm.-Hol. (2 families, 2 genera, 3 species)
Psocidiidae Tillyard 1926 [= Dichentomidae Carpenter 1932]
Dichentomum Tillyard 1926 [= Psocidium, Chaetopsocidium, Metapsocidium, Pentapsocidium, Permentomum, Tillyard 1926; Parapsocidium Zallesky 1937]
tinctum Tillyard 1926 [= D. complexum Carpenter 1926; Psocidium permianum, P. abnormale, P. robustum, P. kansasense, Chaetopsocidium sellardsi, Metopsocidium loxoneurum, Pentapsocidium indistinctum, Permentomum tenuiforme, all Tillyard 1926]
Lophioneuridae Tillyard 1921 [= Cyphoneuridae Carpenter 1932; Zorapsocidae Tillyard 1935]
Cyphoneurodes Bekker-Migdisova 1953
patriciae Beckemeyer 2004
Species No. 2 (not assignable to Elmo taxa, different from above, but may not be well-enough preserved to describe. Collected by Roy Beckemeyer & Joseph Hall, May, 2003.
Caloneurodea (Pterygota: Neoptera: Exopterygota) U.Carb.-Perm. (1 family, 5 genus, 6 species)
Apisdoneuridae Carpenter 1961
Apisdoneura Carpenter 1943
reducta Zimmerman in Tasch and Zimmerman 1962
Uncertain Family:
Gigagramma Bethoux, Nel & Lapeyrie 2004
carpenteri Bethoux, Nel & Lapeyrie 2004
Paleuthygramma Martynov 1930
acuta Carpenter 1943 [per Bethoux, Nel & Lapeyrie 2004]
Anomalogramma Carpenter 1943
parva Carpenter 1943 [per Bethoux, Nel & Lapeyrie 2004]
Uncertain Family
Ligogramma Bethoux, Nel & Lapeyrie 2004
sinuosa Bethoux, Nel & Lapeyrie 2004
wichita Beckemeyer, 2009
Miomoptera (Pterygota: Neoptera: Exopterygota) U.Carb.-Perm. (1 family, 1 genus, 3 species)
Palaeomanteidae Handlirsch 1906 [nom. corr. Palaeomantidae Handlirsch 1906; =Delopteridae Sellards 1909]
Palaeomantis Handlirsch 1904 [= Delopterum Sellards 1909; Pseudodelopterum, Pseudomantis, Leptoneurula, Martynov 1928; Delopsocus Tillyard 1928; Miomantisca, Miomatoneurites, Delopteriella, Zalessky, 1956; Stefanomioptera Guthorl 1962]
minutum (Sellards) 1909 (Delopterum) [= D. dunbari Tillyard 1928]
elongatum (Sellards) 1909 (Delopsocus) [= D. fasciatus Tillyard 1928]
latum (Sellards) 1909 (Delopsocus)
Hemiptera (Pterygota: Neoptera: Exopterygota) Perm.-Hol. (1 family, 2 genera, 2 species)
Homoptera Leach 1815
Archescytinidae Tillyard 1926 [= Permopsyllidae Tillyard 1926; Lithoscyinidae Carpenter 1933; and others]
Paramopsylla Zimmerman in Tasch and Zimmerman 1962
tricubita Zimmerman in Tasch and Zimmerman 1962
Permopsylla Tillyard 1926
Glosselytrodea (Pterygota: Neoptera: Endopterygota) Perm.-Jur. (1 family, 1 genus, 1 species)
Permoberothidae Tillyard 1932
Permoberotha Tillyard 1932 [= Dictyobiella Tillyard 1937]
villosa Tillyard 1932 [ = P. convergens Tillyard 1937; D. nervosa Tillyard, 1937]
Mecoptera (Pterygota: Neoptera: Endopterygota) Perm.-Holo. (2 family, 2 genera, 2 species) [Order mentioned by Tasch & Zimmerman as having been found at Midco, but they listed no taxa names.]
Permopanorpidae Tillyard 1926
Permopanorpa Tillyard 1926
inaequalis Tillyard 1926
Permochoristidae Tillyard 1918
Undetermined genus (Specimen collected by Tasch now in RJB collection and yet to be identified)
Undetermined species (Specimen collected by Tasch now in RJB collection and yet to be identified))
Coleoptera (Pterygota: Neoptera: Endopterygota) Perm.-Holo. (1 family, 1 genus, 1 species)
Undetermined Family [Although Ponomarenko & Kirejtshuk (2007) placed Permocoleus in Tschekardocoleidae Rhodendorf, 1944]
Permocoleus Lubkin & Engel 2005
wellingtonensis Lubkin & Engel 2005
Uncertain Order (Pterygota: Neoptera: Polyneoptera: Dictyoptera (Stem-group Mantodean)) (1 family, 1 genus, 1 species)
Strephocladidae Martynov 1938
Homocladus Carpenter 1966 [= Paracladus Carpenter, 1966, per Bethoux, Beckemeyer, Engel and Hall, 2010]
grandis Carpenter 1966 [= H. ornatus Carpenter, 1966 = P. retardatus Carpenter, 1966 per Bethoux, Beckemeyer, Engel, and Hall, 2010]
Summary:
Apterygota: 1 Order; 1 Family; 1 Genus; 1 Species
Palaeoptera: 6 Orders; 13 Families; 19 Genera; 45 Species
Neoptera: 12 Orders; 22 Families; 34 Genera; 38 Species
Total: 19 Orders; 36 Families; 54 Genera; 84 Species
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