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Liomopterum ornatum specimen currently in the collection of Roy J. BeckemeyerMIDCO, OKLAHOMA - THE FIRST OKLAHOMA FOSSIL INSECT LAGERSTATTEN

A Web Page by Roy J. Beckemeyer (send mail to royb at southwind.net)
Last updated: 1 June 2005

The image on the right is a scanned (1200 dpi) fossil fore wing of a Protorthopteran: Liomopteridae: Liomopterum ornatum Sellards 1909.  ex Tasch collection (from discarded bulk material collected by Dr. Paul Tasch from his Noble VII site, Midco Insect Bed, Noble County, Oklahoma).  Specimen RJB001b, identified by Roy J. Beckemeyer, March, 2000.  Portion of fore wing.  The shell-shaped object in the upper right is a conchostracan (clam shrimp).  


An Ongoing Narrative About The Midco Fossil Insect BEds...

The Midco, Oklahoma fossil beds were discovered by G.O. Raasch in Noble County, Oklahoma.  In 1939 he sent about seventy fossil insect specimens he had collected in Noble County to Frank Carpenter.  The following summer Carpenter and Raasch collected over 5,000 "well-preserved" insect fossils. 

This locality is about 140 miles from the Elmo site in Kansas and is also part of the Wellington formation (the Wellington Sea covered parts of Oklahoma, Kansas, Nebraska, and Colorado).  The insect-bearing sediments at Midco were apparently laid down in a playa, Midco Salt Lake, which held only algae and Conchostraca (Crustacea).  The insects were apparently carried into the lake by flooding of small streams.  Plants did not grow near the lake.  This site was thus quite different in nature from the Elmo location.


A Midco, Oklahoma fossil insect site near Perry, OKAnother difference from Elmo is the widespread character of the Midco deposits: insect-bearing Midco rocks are spread over an area of some 400 square miles while Elmo is a small site.

[The photograph on the left was taken by Roy J. Beckemeyer at a Midco site near Perry, Oklahoma on 16 May, 2000.  The arrows point to two of the insect layers of Raasch's Midco deposit.  A number of plant and insect fossils as well as many conchostracans were found in rock slabs from this site.]

Carpenter returned to the Midco site in the years from 1948 to 1957.  The Harvard collection totaled about 8,000 specimens at the time of Carpenter's last publication on the Midco insect fauna.  (See Carpenter 1947 and 1979 in the bibliography for a description of the collecting sites and fossils.)  At the end of his first look at the Oklahoma fossils in 1947, Carpenter had found it necessary to study "as many as possible of the Palaeozoic insects already described from European and North American deposits before continuing with the new material" (Carpenter, 1979).  As a result, his two papers covered only the Palaeoptera and he did not include any Neoptera in either of the two papers.  
Fragment of a protodonate wing found at a Midco site in May, 2000
[The picture on the left is a scanned image of a fragment of a large wing that appears to be that of a protodonate, likely a Meganeuropsis. (M. permiana was described from the Elmo Kansas site, and M. americana from the Midco site.)   Detailed study is pending, but the size and nature of the venation is indicative of this preliminary diagnosis.  The fragment was collected at the site pictured above by Roy J. Beckemeyer in May, 2000.]

In the late 1950's and early 1960's, Dr. Paul Tasch of the University of Wichita (now Wichita State University) began a study of the palaeolimnology and particularly the conchostracan-bearing beds of the Kansas and Oklahoma Permian (Tasch, 1958, 1961, 1962, 1963, Tasch & Zimmerman, 1959, 1962).  He came across many insect bearing strata and collected extensively in Raasch's Midco Beds as well as other newly discovered beds in Oklahoma (Noble and Kay counties) and Kansas (particularly Sumner County).  He attempted to correlate the Elmo and Midco with the other insect bearing strata and to trace the layers through the two-state region.


Tasch and Zimmerman (1962) surveyed some of the Permian insect fauna represented in their collections.  They included both Palaeoptera and Neoptera, but did not publish on any taxa on which Carpenter had yet to publish, stating that "...the systematics given in this paper far from exhaust material still being processed and studied."  

Chelopterum peregrinum, a fossil wing in the collection of Roy BeckemeyerThe image on the right is a scanned (1200 dpi) fossil Protorthopteran: Chelopteridae: Chelopterum peregrinum Carpenter 1950.  Originally described from the Elmo, Kansas beds.  Specimen is ex Tasch collection (see notes at the top of this page).  From Noble IIA, Bed 14, collected by Paul Tasch.  Specimen No. RJB002, portion of fore wing, identified March, 2000 by Roy J. Beckemeyer. 

I am not aware of any further papers by either Zimmerman or Tasch on the insect fauna of these beds, but need to review both the specimens in the Tasch collection in the Geology Dept. at Wichita State University and the Tasch papers in the Special Collections Library at the University.  Carpenter (1992) failed to include reference to any of the new taxa described by Zimmerman in Tasch & Zimmerman (1962), but I have included them on the checklist of Midco Permian Insects.

I recently met with Larry Skelton, manager of the Kansas Geological Survey Well Sample Library in Wichita to discuss the Kansas Permian deposits.  He had "rescued" some of the Tasch collection bulk material from Kansas and Oklahoma when it had been readied for disposal when the Geology Department was moving.   He transferred to me some 6 boxes of material, labeled as to site, which I am going over in search of additional insect material.  Thus far, I have come across wings of two species, both Protorthoptera:  Liomopterum ornatum Sellards 1909 (which had been listed by Tasch and Zimmerman) and Chelopterum peregrinum Carpenter 1950 (which had not been listed by them).  I am continuing to work over the material and will report findings here.  


Left: A drawing of C. peregrinum by Roy Beckemeyer based on a reconstruction by Frank Carpenter.  Colors are of course imaginary.

 

 


In May of 2000 I visited Noble County and was able to collect a small amount of material from one of Raasch's localities.  Fossils found from the slabs to date include the possible protodonate wing fragment pictured above, and a number of smaller wings which are also being studied.  One of the smaller wings appears on first look to be that of an adult mayfly,  family Protereismatidae.   -- Roy J. Beckemeyer, 22 May, 2000.

 

Protereisma species - roughly 40% of hind wing - collected from Raasch site 9 by Roy Beckemeyer, May, 2000Left is a picture of the mayfly wing.  It is a hind wing of an adult Protereismatidae: Protereisma sp.  It is not P. directum Carpenter 1979, nor is it P. latum Sellards 1907, which Tasch and Zimmerman reported from Midco.  It appears somewhat similar to P. permianum Sellards 1907, but may be an undescribed species.  I hope to compare it to the series of material discussed by Tillyard and Carpenter in their reviews of the Elmo Ephemeroptera, and to the undescribed adult Ephemeroptera wings from Midco that Carpenter mentioned in his 1979 paper. - Roy Beckemeyer, April, 2003.  NOTE: Since this appears to be a new species I intend to publish a description soon, so please do not reproduce this figure. Roy Beckemeyer, May 2003.

 

 

 

 

In late 2002 and early 2003, Joseph Hall, a graduate student in geology at Wichita State University studying the stratigraphy and sedimentology of the Wellington Formation in Kansas and Oklahoma, and I visited a number of insect localities in Oklahoma and Kansas.  The Delopterum latum wing shown at left was collected by Joseph on a trip he made to Raasch's Site 5 in Noble County in March, 2003.  - Roy Beckemeyer, April, 2003.

 

 

 

 


 

Tasch II-A (Raasch Station 3), Noble Co., OklahomaClose-up view of insect layer at Tasch II-A

Left: A locality visited by Roy Beckemeyer and Joseph Hall in April and May, 2003: Tasch II-A (Raasch 3), in Noble County, Oklahoma.  One fossil insect layer was worked and yielded a number of small wings and wing fragments.  Pictures will be added here soon.  Right: A close-up view of the location of the insect bed within a unit of alternating shales and dolomite. 


New Martynovia species: The pictures below are counterpart (left, the impression of the wing) and part (right, the dorsal surface of the wing) of an exciting specimen from Raasch 9 collected by Roy Beckemeyer and Joseph Hall on 1 May, 2003.  The counterpart constitutes nearly the entire wing, the part preserves about 2/3 of the wing in two fragments (the distal portion a nearly paper-thin piece that fractured off the substrate during exposure of the fossil).  The fossil is a well-preserved specimen of the order Diaphanopterodea, family Martynoviidae.  Once known only from the Wellington of Kansas and Oklahoma, Bethoux, Nel, Lapeyrie, Gand, and Galtier (2003) recently described several taxa from this family that were found in the Salagou Formation in France.  The specimen below has the wing basal venation very well preserved, and is of some significance.  It is probably a new species (apparently of genus Martynovia, and similar to M. protohymenoides Tillyard, 1932, from Elmo) and I will be publishing a description soon.  (There are 2 species of Martynovia known from Elmo, and one from Midco - this may be the second Midco species.)  Please do not reproduce these figures.  UPDATE:  June, 2004.  New species description now published.  See "A new species of Martynovia Tillyard, 1932 (Insecta: Daphanopterodea: Martynoviidae) from the Lower Permian Welllington Formation of Noble County, Oklahoma", by Roy J. Beckemeyer, Journal of the Kansas Entomological Society 77(2):127-131 (28 April 2004), which names this fossil insect "Martynovia halli Beckemeyer, 2004" - named for Joseph Hall.

Beckemeyer/Hall specimen of Daphanopterodea wing from Noble CO. OKCounterpart of Beckemeyer/Hall specimen of Martynoviidae wing from Noble Co., OK

 

 

 

 

 

 

 


New Specimen of Oligotypus tillyardi collected by Joseph Hall and Michael Montgomery, 31 July 2003 - Link here for photos


Chelopterum peregrinum from OklahomaLeft: A specimen of Protorthoptera: Chelopteridae: Chelopterum peregrinum collected by Joseph Hall from Raasch Locality 4, Noble County, Oklahoma in August, 2003.  

Link here to more photos of the fossils on the slab that contained this specimen: CHELOPTERIDAE/PROBNIDAE

 

 


LINK HERE for a page of photos of Protorthoptera: Probnidae: Probnis cf. speciosa Tillyard 1939, including a rather spectacular set of four wings collected by Joseph Hall in Sept. 2003.


LINK HERE for a page of photos of a specimen of an Odonatoptera (Protanisoptera): Ditaxineuridae: Ditaxineura sp. taken from the Billings 3 locality, Sept. 2003 by Joseph Hall and Michael Montgomery.


LINK HERE for a page of photos of a specimen of the mayfly Ephemeroptera: Protereismatidae: Proteriesma directum Carpenter, 1979, taken from locality Noble IIA, 19 July, 2003 by Joseph Hall.


Link Here for a page with a species new to Midco, described by Carpenter from Elmo, Kansas: Stereopterum rotundum, collected from Billings 3, Noble County, by Roy Beckemeyer.


Link Here for fantastic images of a nearly complete specimen of Glaphyrophlebia cf ovata, with articulated wings, head, thorax, legs and abdomen, collected in April, 2004 from locality Raasch 5 by Joseph Hall.


Link Here for more fantastic images of a nearly complete specimen of Misthodotes cf edmundsi Carpenter, 1979, with portions of four wings, abdomen, complete thorax, collected by Joseph Hall from Billings 3 locality 29 April, 2004.


Link Here for image of a scorpionfly wing, Permopanorpa inaequalis Tillyard, 1926.  Although Tasch & Zimmerman 1962 mentioned having taken Mecoptera from Midco, they did not list or illustrate any taxa, so this is the first specific information provided for Mecoptera of the Oklahoma Permian.

 


Link Here for images of a Caloneurodea: Paleuthygramma acuta Carpenter, 1943, comprising both forewings, part of thorax, and a leg fragment, collected in Sept. 2004 by Bill May at Billings 3.

 


LINK Here for images of a Protelytroptera elytron from Billings 3 collected by Joseph hall in Sept. 2004.


LINK HERE for images of a fore wing of the Protorhtoptera: (sensu Carpenter, 1992) or Grylloblattida: Lemmatophorina: (sensu Rasnitsyn & Quicke, 2002) Paraprisca fragilis (Sellards) 1909 (Family Lemmatophoridae).  The species was described from the Elmo deposits and is newly recorded here for the Midco.  Collected by Joseph Hall in 2004 and identified and photographed by ROy Beckemeyer in May, 2005.


SOME NON-INSECT PERMIAN FOSSILS FROM NOBLE COUNTY:


plantR9.JPG (69893 bytes)Left:  A plant fossil from Raasch 9 locality collected by Roy Beckemeyer, 16 May, 2000.  

 

 


Lungfish tooth plate from vertebrate layer at Billings 3 Locality - collected by Joseph Hall

Left:  A lungfish tooth plate from the vertebrate bed at Billings 3 in Noble County, Oklahoma.  Collected in Sept. 2004 by Joseph Hall.    

 

 

 

 

 


 

The Paleontologists who worked the Midco site:

 


Insects described from Midco:

Midco Fossil Insect Checklist


LINKS TO OTHER OKLAHOMA PALEONTOLOGY WEB SITES


 

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