OUTLINE OF CLASSIFICATION OF THE SUPERCLASS HEXAPODA:
Based on the outline on p. 504 of Carpenter's "Treatise on Invertebrate Paleontology. Part R. Arthropoda 4. Volume 4. Hexapoda.", Geol. Soc. Am. & Univ. Kansas, 1992, with added data on the Kansas Elmo fossils, and with English names for the taxa based on D.J. Borror, D.M. DeLong, & C.A. Triplehorn, "An Introduction to the Study of Insects" (4th ed.), Holt, Rinehart & Winston, 1976, and on CSIRO's "The Insects of Australia" 2nd edition.
Prepared by Roy J. Beckemeyer.
Key: Level TAXON Nr. of fossil genera/Nr. of Elmo fossil genera Geological eras of occurrence (Magenta if at Elmo)
Superclass HEXAPODA 5,188 Devonian-Holocene
Class and Order COLLEMBOLA (Springtails) 14 Devonian-Holocene
Class and Order PROTURA (Proturans) 0 Holocene
Class and Order DIPLURA (Diplurans) 4 Paleocene-Holocene
Class INSECTA (Insects) 5,169/93 Upper Carboniferous-Holocene
Subclass APTERYGOTA (primitively wingless insects) 8/1 Upper Carboniferous-Holocene
Order ARCHAEOGNATHA (Bristletails) 5/1 Upper Carboniferous-Holocene
Order ZYGENTOMA (Silverfish) 3 Oligocene-Holocene
Subclass PTERYGOTA (winged and secondarily wingless insects) 5,160/92 Upper Carboniferous-Holocene
Infraclass PALAEOPTERA (wings cannot be folded flat against dorsum of abdomen) 502/20 Upper Carboniferous-Holocene
Order EPHEMEROPTERA (Mayflies) 59/2 Upper Carboniferous-Holocene
Order PALAEODICTYOPTERA (wing-like prothoacic lobes, sucking mouthparts) 125/4 Upper Carboniferous-Permian
Order MEGASECOPTERA (similar to previous, but with narrower wing bases) 34/2 Upper Carboniferous-Permian
Order DIAPHANOPTERODEA (similar to above but could fold wings, but differently from the modern Neoptera) 20/3 Upper Carboniferous-Permian
Order PROTODONATA (large to gigantic Dragonfly precursors) 13/4 Upper Carboniferous-Permian
Order ODONATA 204/4 (Dragonflies and Damselflies) Upper Carboniferous-Holocene
Order UNCERTAIN, Infraclass PALAEOPTERA 47/1
Infraclass NEOPTERA (wings can be folded flat against dorsum of abdomen) 4,656/72 Upper Carboniferous-Holocene
Division EXOPTERYGOTA (NEOPTERA with simple metamorphosis) 1,393/57 Upper Carboniferous-Holocene
Order PERLARIA (Most authors use the modern name PLECOPTERA: Stoneflies) 28 Permian-Holocene
Order PROTORTHOPTERA (Orthopteroids, many Stonefly-like, probably not a natural group) 236/24 Upper Carboniferous-Permian
Order BLATTARIA (Cockroaches) (Not covered in Treatise)/2 Upper Carboniferous-Holocene
Order ISOPTERA (Termites) 33 Cretaceous-Holocene
Order MANTODEA (Praying Mantids) 4 Oligocene-Holocene
Order PROTELYTROPTERA (small insects related to Cockroaches) 236/9 Permian-Cretaceous
Order DERMAPTERA (Earwigs) 16 Jurassic-Holocene
Order ORTHOPTERA (Grasshoppers, Crickets & their relatives) 166/5 Upper Carboniferous-Holocene
Order GRYLLOBLATTODEA (Rock Crawlers) 0 Holocene
Order TITANOPTERA (Orthopteroid insects of moderate to large size) 8 Triassic
Order PHASMATODEA (Walking Sticks) 13 Triassic-Holocene
Order EMBIOPTERA (Webspinners) 5 Oligocene-Holocene
Order PSOCOPTERA (Psocids - "booklice", "barklice") 58/7 Permian-Holocene
Order ZORAPTERA (Zorapterans) 0 Holocene
Order MALLOPHAGA (Chewing Lice) 0 Holocene
Order ANOPLURA (Sucking Lice) 0 Holocene
Order CALONEURODEA (Orthopteroid-like insects) 16/5 Upper Carboniferous-Permian
Order MIOMOPTERA (small insects with biting mouthparts and nearly equal wings) 10/1 Upper Carboniferous-Permian
Order THYSANOPTERA (Thrips) 43 Permian-Holocene
Order HEMIPTERA (True Bugs) 728/4 Permian-Holocene
Division ENDOPTERYGOTA (NEOPTERA with complete metamorphosis) 3,203/7 Permian-Holocene
Order COLEOPTERA (Beetles) 1,397 Permian-Holocene
Order STREPSIPTERA (Twisted-wing insects) 4 Oligocene-Holocene
Order NEUROPTERA (Net-winged insects) 125 Permian-Holocene
Order GLOSSELYTRODEA (small insects with elytriform wings) 11/1 Permian-Jurassic
Order TRICHOPTERA (Caddisflies) 112 Permian-Holocene
Order LEPIDOPTERA (Butterflies and Moths) 111 Cretaceous-Holocene
Order MECOPTERA (Scorpionflies) 82/6 Permian-Holocene
Order SIPHANOPTERA (Fleas) 1 Oligocene-Holocene
Order DIPTERA (True Flies) 708 Triassic-Holocene
Order HYMENOPTERA (Ants, Wasps, Bees) 652 Triassic-Holocene
Order UNCERTAIN, Infraclass NEOPTERA 60/8
Subclass PTERYGOTA, Order UNCERTAIN 2
Class INSECTA, Subclass UNCERTAIN 1
Superclass HEXAPODA, Class UNCERTAIN 1
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