CHECKLIST OF ELMO, KANSAS PERMIAN INSECTS
Orders in taxonomic sequence per
Carpenter (1992):
"Treatise
on Invertebrate Paleontology. Part R. Arthropoda 4. Vol. 3 & 4. Hexapoda,"
Taxonomic assignments and synonymies per that reference
as well.
Synonyms given when they relate to names used in the works of
Sellards, Carpenter, and Tillyard.
For other synonyms, see the Treatise
volumes.
A Web Page by Roy J. Beckemeyer
Last updated: 7 August 2007
Revised 22/7/07: Added taxa described by Béthoux & Beckemeyer 2007. Revised 4/8/00: Added Meganeuridae: Megatypus parvulus Engel 1998, a taxon missed from the original compilaiton. This species brings the Protodonata species count to 8 and the total species count to 150. Revised 2/12/00: Corrected typographic errors: Misthodotes ovatus Tillyard 1932 becomes Misthodotes ovalis Tillyard 1932 (Ephemeroptera); date of description of Probnis speciosa Tillyard changed from 1939 to 1937 (Protorthoptera); date of description of Palaeomantis minutum (Sellards) changed from 1904 to 1909 (Miomoptera); Palaeomantis elongates becomes Palaeomantis elongatum and Palaeomantis latus becomes Palaeomantis latum (Miomoptera).
Added link to page of images of Protodonata: Tupus permianus Sellards 1906. Removed Tupus vetustus and made it a synonym of T. permianus per Carpenter, which I had misinterpreted originally. Reduces number of species to 149.
Added Oligotypus sp. Carpenter 1931 which brough species number back to 150. Corrected spelling of M. parvus Engel (Protodonata) from M. parvulus. [3 March 2002].
Added link to
page of images of Oligotypus tillyardi Carpenter 1931 - a specimen was
collected from the Midco, Oklahoma beds in August 2003 by Michael Montgoemry and
Joseph Hall.
Use your web browser "Find" function to search the page for known taxa.
Key:
No indentation:
Order (Higher Taxon) Geological time of occurrence (Number of genera,
species)
One tab: Suborder
Author Year described [synonymies in brackets]
Two tabs: Family Author Year
described [synonymies in brackets]
Three tabs: Genus Author Year
described [synonymies in brackets]
Four tabs: species Author (in parentheses
if described in another Genus) Year described (Original
Genus)
[synonymies in brackets]
Archaeognatha (Apterygota) U.Carb.-Hol. (1 genus, 1 species)
Dasyleptidae Sharov 1957
Lepidodasypus Durden 1978
sharovi Durden 1978
Ephemeroptera (Pterygota: Palaeoptera) U.Carb.-Hol. (2 genera, 8 species) [=Plectoptera])
Misthodotidae Tillyard 1932
Misthodotes Sellards 1909 [nom. subst. pro Dromeus Sellards 1907;= Eudoter Tillyard 1936]
obtusus (Sellards) 1909 (Dromeus)
biguttatus Tillyard 1932
delicatulus (Tillyard) 1932 (Eudoter)
ovalis Tillyard 1932
Protereismatidae Lameere 1917 [= Protereismopheridae Sellards 1907]
Protereisma Sellards 1907 [= Protechma, Prodromus, Bantiska, Pinctodia, Rekter Sellards, 1907; Esca, Mecus Sellards 1909; Loxophlebia Martynov 1928]
permianum Sellards 1907 [= Protechma acuminatum Sellards 1907; Prodromus rectus Sellards 1907; Scopus gracilis Sellards 1907; Protereisma sellardsi Tillyard 1932, P. gracile Sellards 1907, P. minus Sellards 1907]
elongatum (Sellards) 1907 (Bantiska)
arcuatum (Sellards) 1907 (Rekter)
latum Sellards 1907
insigne Tillyard 1932
Palaeodictyoptera (Pterygota: Palaeoptera) U.Carb.-Perm. (4 genera, 4 species)
Calvertiellidae Martynov 1931
Calvertiella Tillyard 1925
permiana Tillyard 1925
Elmoboriidae Carpenter 1976
Elmoboria Carpenter 1976
piperi Carpenter 1976
Spilapteridae Handlirsch 1906
Dunbaria Tillyard in Dunbar & Tillyard 1924
fasciipennis Tillyard in Dunbar & Tillyard 1924
Uncertain Family
Kansasia Tillyard 1937
pulchra Tillyard 1937
Megasecoptera (Pterygota: Palaeoptera) U.Carb.-Perm. (2 genera, 5 species) [= Protohymenoptera Tillyard 1924]
Protohymenidae Tillyard 1924
Protohymen Tillyard 1924 [= Pseudohymen Martynov 1932, Pseudohymenopsis Zalessky 1956]
permianus Tillyard 1924
elongatus Carpenter 1930
readi Carpenter 1933
tenuis Carpenter 1938
Permohymen Tillyard 1924
schucherti Tillyard 1924
Diaphanopterodea
(Pterygota: Palaeoptera) U.Carb.-Perm. (3 genera, 6 species)
Asthenohymenidae Tillyard 1924
Asthenohymen Tillyard 1924 [= Karoohymen Riek 1976]
dunbari Tillyard 1924 [= A. gracilis, A. kansasensis, A. stenobasis, Tillyard, 1926; Doter minor Carpenter 1930]
affinis Tillyard 1926 [= A. stigmatizans Tillyard 1926; Doter affinis Carpenter 1930]
pusillus Tillyard 1926 [= Doter pusillus Carpenter 1930]
Elmoidae Tillyard 1937
Elmoa Tillyard 1937
trisecta Tillyard 1937
Martynoviidae Tillyard 1932 [= Martynoviella Tillyard 1932]
Martynovia Tillyard 1932
insignis Tillyard 1932
protohymeniodes(Tillyard) 1932 (Martynoviella)
Protodonata (Pterygota: Palaeoptera) U.Carb.-Perm. (4 genera, 8 species) [= Meganisoptera Martynov 1932]
Meganeuridae Handlirsch 1906
Meganeuropsis Carpenter 1939
permiana Carpenter 1939
Megatypus Tillyard 1925
parvus Engel 1998
schucherti Tillyard 1925
Tupus Sellards 1906 [= Typus Sellards 1909, unjustified emendation; Meganeurela Handlirsch 1906 among others]
permianus Sellards 1906 [= Megatypus vetustus (Carpenter) 1933]
readi Carpenter 1933
Paralogidae Handlirsch 1906
Oligotypus Carpenter 1931
sp. Carpenter 1931 [larger than tillyardi, but deemed by Carpenter to be insufficient material for defining a species]
Odonata (Pterygota: Palaeoptera) Perm.-Hol. (4 genera, 7 species)
Archisozygoptera Handlirsch 1906 [= Protozygoptera Tillyard 1925]
Kennedyidae Tillyard 1925
Kennedya Tillyard 1925
mirabilis Tillyard 1925
tillyardi Carpenter 1939
reducta Capenter 1939
Progoneura Carpenter 1931
minuta Carpenter 1931
Protanisoptera Carpenter 1931
Ditaxineuridae Tillyard 1926
Ditaxineura Tillyard 1926
anomalostigma Tillyard 1926
celulosa Carpenter 1933
Uncertain Suborder
Uncertain Family
Campotaxineura Tillyard 1937
ephialtes Tillyard 1937
Uncertain Order (Pterygota: Palaeoptera) (1 genus, 1 species)
Uncertain Family
Permoneura Carpenter 1931
lameerei Carpenter 1931
Protorthoptera (Pterygota: Neoptera: Exopterygota) U.Carb.-Triass. (26 genera, 36 species) [= Protoperlaria Tillyard 1928; Protoblattoidea Handlirsch 1906, Paraplecoptera Martynov 1938, among others]
Blattinopsidae Bolton 1925
Blattinopsis Giebel 1867 [= Oryctoblattina Scudder 1879 among others]
kukalovae Carpenter 1966
Glaphyrophlebia Handlirsch 1906 [= Pursa, Sindon, Sellards, 1909]
speciosa (Sellards) 1909 (Sindon)
ovata (Sellards) 1909 (Pursa)
Chelopteridae Carpenter 1950
Chelopterum Carpenter 1950
peregrinum Carpenter 1950
Demopteridae Carpenter 1950
Demopterum Carpenter 1950
gracile Carpenter 1950
Heteroptilidae Carpenter 1976
Heteroptilon Carpenter 1976
costale Carpenter 1976
Lemmatophoridae Sellards 1909 [= Ortaidae Sellards 1909, Lecoriidae handlirsch 1919]
Lemmatophora Sellards 1909
typa Sellards 1909 [= typica Tillyard 1928; anomala, elongata, delicosa, hirsuta, Sellards 1909]
Artinska Sellards 1909 [= Estadia, Lectrum, Orta, Sellards 1909]
clara Sellards 1909 [= medialis, gracilis, pecta, major, extensa, Sellards, 1909; Estadia elongata, arcuata, tenuis, Sellards, 1909; Estadia tripunctata Carpenter 1926; Lectrum anomalum Sellards, 1909]
sellardsi Tillyard 1928
ovata (Sellards) 1909 (Orta)
Lisca Sellards 1909
minuta Sellards 1909 [= Lemmatophora reducta, minuta, Tillyard 1928; Lemmatophora cubitalis Handlirsch 1937]
Paraprisca Handlirsch 1919 [nom. subst. pro Prisca Sellards 1909]
fragilis (Sellards) 1909 (Prisca)
grandis Carpenter 1935 [= P. stigmatica Tillyard 1937]
Lecorium Sellards 1909 [= Stemma Sellard 1909, Sellardsia Tillyard 1928, among others]
elongatum Sellards 1909 [= Stemma elegans, extensa, Sellards, 1909; Sellardsia kansasensis, lecorioides Tillyard, 1928]
Liomopteridae Sellards 1909
Liomopterum Sellards 1909 [= Horates Sellards 1909]
ornatum Sellards 1909 [= L. extensum Sellards 1909]
elongatum (Sellards) 1909 (Horates)
sellardsi Carpenter 1950
Semopterum Carpenter 1950
venosum Carpenter 1950
Tapopterum Carpenter 1950
celsum Carpenter 1950
Phenopteridae Carpenter 1950
Phenopterum Carpenter 1950 [nom. subst. pro. Lepium Sellards 1909]
elongatum (Sellards) 1909 (Lepium) [= Lepium reticulatum Sellards 1909; Lepium sellardsi Handlirsch 1919]
Probnidae Sellards 1909 [nom. correct. pro Probnisidae Sellards 1909]
Probnis Sellards 1909 [= Espira, Stoichus, Stinus, Sellards 1909]
speciosa Tillyard 1937 [= P. coriacea, Espira obscura, Stoichus elegans, S. arcuatus, S. minor, S. tenuis, Stinus brevi-cubitalis, Sellards 1909; Probnis sellardsi Handlirsch 1919]
Protembiidae Tillyard 1937
Protembia Tillyard 1937 [= Telactinopteryx Tillyard 1937]
permiana Tillyard 1937 [= Telactinopteryx striatipennis Tillyard 1937]
Psoropteridae Carpenter 1976
Psoroptera Carpenter 1976
cubitalia Carpenter 1976
Steropteridae Carpenter 1950
Steropterum Carpenter 1950
rotundum Carpenter 1950
maculosum Carpenter 1966
breve Carpenter 1966
Strephocladidae Martynov 1938
Homocladus Carpenter 1966
grandis Carpenter 1966
ornatus Carpenter 1966
Paracladus Carpenter 1966
retardatus Carpenter 1966
Tococladidae Carpenter 1966
Tococladus Carpenter 1966
rallus Carpenter 1966
Opisthocladius Carpenter 1976
arcuatus Carpenter 1976
strictus Carpenter 1976
Uncertain Family
Aibolitus Béthoux & Beckemeyer 2007
minutus Béthoux & Beckemeyer 2007
Atava Sellards 1909
ovata Sellards 1909
Elmopterum Béthoux & Beckemeyer 2007
rotundum Béthoux & Beckemeyer 2007
Lecopterum Sellards 1909
delicosum Sellards 1909
Blattaria (Pterygota: Neoptera: Exopterygota) U.Carb.-Hol. (2 genera, 4 species)
Spiloblattinidae Handlirsch
Permoblattina Tillyard 1937
curta (Sellards) 1908 (Etoblattina)
permiana (Sellards) 1908 (Etoblattina)
Pycnoblattina Sellards 1908
compacta (Sellards) 1908 (Puknoblattina)
curvata (Sellards) 1908 (Puknoblattina)
Protelytroptera (Pterygota: Neoptera: Exopterygota) Perm.-Cret. (9 genera, 15 species)
Archelytridae Carpenter 1933
Archelytron Carpenter 1933
superbum Carpenter 1933
Elytroneuridae Carpenter 1933
Elytroneura Carpenter 1933
permiana Carpenter 1933
Megelytridae Carpenter 1933
Megelytron Tillyard 1931
robustum Tillyard 1931
Permelytridae Tillyard 1931 [= Blattelytridae Tillyard 1931; Acosmelytridae Tillyard 1931]
Permelytron Tillyard 1931
schucherti Tillyard 1931
Blattelytron Tillyard 1931
permianum Tillyard 1931
Parablattelytron Tillyard 1931 [= Acosmelytron Tillyard 1931]
subincisum Tillyard 1931
rectum Tillyard 1931
elongatum (Tillyard) 1931 (Acosmelytron) [= P. acutum (Tillyard) 1931 (Acosmelytron)
delicatum (Tillyard) 1931 (Acosmelytron)
latum Carpenter 1937
Protelytridae Tillyard 1931
Protelytron Tillyard 1931
permianum Tillyard 1931 [= tillyardi Carpenter 1933]
angustum Carpenter 1933
furcatum Carpenter 1939
Permelytropsis Carpenter 1933
cubitalis Carpenter 1933
Uncertain Family
Protelytropsis Tillayrd 1931
grandis Tillyard 1931
Orthoptera (Pterygota: Neoptera: Exopterygota) U.Carb.-Hol. (6 genera, 8 species)
Oedischiidae Handlirsch 1906
Paroedischia Carpenter 1966
rectu Carpenter 1966
maculata Carpenter 1966
Permelcanidae Sharov 1962
Promartynovia Tillyard 1937
venicosta Tillyard 1937
Anelcanidae Carpenter 1987 [nom. subst. pro Parelcanidae Carpenter 1966]
Anelcana Carpenter 1987[nom. subst. pro Parelcana Carpenter 1966]
dilatata (Carpenter) 1966 (Parelcana)
Petrelcana Carpenter 1966
elongata Carpenter 1966
Permoraphididae Tillyard 1932
Permoraphidia Tillyard 1932
americana Tillyard 1932
grandis Carpenter 1943
Uncertain family
Crinoedischia Béthoux & Beckemeyer 2007
buttsi Béthoux & Beckemeyer 2007
Psocoptera (Pterygota:
Neoptera: Exopterygota) Perm.-Hol. (7 genera, 12 species)
Psocidiidae Tillyard 1926 [= Dichentomidae Carpenter 1932]
Dichentomum Tillyard 1926 [= Psocidium, Chaetopsocidium, Metapsocidium, Pentapsocidium, Permentomum, Tillyard 1926; Parapsocidium Zallesky 1937]
tinctum Tillyard 1926 [= D. complexum Carpenter 1926; Psocidium permianum, P. abnormale, P. robustum, P. kansasense, Chaetopsocidium sellardsi, Metopsocidium loxoneurum, Pentapsocidium indistinctum, Permentomum tenuiforme, all Tillyard 1926]
minimum (Tillyard) 1926 (Psocidium)
latum Carpenter 1932
parvulum Carpenter 1932
grande Carpenter 1933
Permopsocidae Tillyard 1926
Prognopsocus Tillyard 1926
permianus Tillyard 1926 [= P. pusillus Tillyard 1926]
Permopsocus Tillyard 1926 [= Ancylopsocus Tillyard 1926]
latipennis Tillyard 1926 [= P. congener, P. enderleini Tillyard 1926, Ancylopsocus insolitus Tillyard 1926]
ovatus Carpenter 1939
Lithopsocidium Carpenter 1932
permianum Carpenter 1932
Orthopsocus Carpenter 1932
simgularis Carpenter 1932
Lophioneuridae Tillyard 1921 [= Cyphoneuridae Carpenter 1932; Zorapsocidae Tillyard 1935]
Cyphoneura Carpenter 1932
permiana Carpenter 1932
Cyphoneurodes Becker-Migsisova 1953
reducta (Carpenter) 1932 (Cyphoneura)
Caloneurodea (Pterygota: Neoptera: Exopterygota) U.Carb.-Perm. (5 genera, 6 species)
Anomalogrammatidae Carpenter 1943 [nom. subst. pro Anomalogrammidae Carpenter 1943]
Anomalogramma Carpenter 1943
parvum Carpenter 1943
Apisdoneuridae Carpenter 1961
Apisdoneura Carpenter 1943
flexa Carpenter 1943
Paleuthygrammatidae Carpenter 1943[nom. subst. pro Paleuthygrammidae Carpenter 1943]
Paleuthygramma Martynov 1930
acutum Carpenter 1943
Permobiellidae Tillyard 1937
Permobiella Tillyard 1937
perspicua Tillyard 1937
Pleisiogrammatidae Carpenter 1943 [nom. subst. pro Plesiogrammidae Carpenter 1943]
Pleisiogramma Carpenter 1943
mediale Carpenter 1943 [=reducta Carpenter 1943]
Miomoptera (Pterygota: Neoptera: Exopterygota) U.Carb.-Perm. (1 genus, 4 species)
Palaeomanteidae Handlirsch 1906 [nom. corr. Palaeomantidae Handlirsch 1906; =Delopteridae Sellards 1909]
Palaeomantis Handlirsch 1904 [= Delopterum Sellards 1909; Pseudodelopterum, Pseudomantis, Leptoneurula, Martynov 1928; Delopsocus Tillyard 1928; Miomantisca, Miomatoneurites, Delopteriella, Zalessky, 1956; Stefanomioptera Guthorl 1962]
minutum (Sellards) 1909 (Delopterum) [= D. dunbari Tillyard 1928]
elongatum (Sellards) 1909 (Delopsocus) [= D. fasciatus Tillyard 1928]
latum (Sellards) 1909 (Delopsocus)
kansasum (Carpenter) 1939 (Delopterum)
Hemiptera (Pterygota: Neoptera: Exopterygota) Perm.-Hol. (4 genera, 10 species)
Homoptera Leach 1815
Archescytinidae Tillyard 1926 [= Permopsyllidae Tillyard 1926; Lithoscyinidae Carpenter 1933; and others]
Archyesctina Tillyard 1926
permiana Tillyard 1926
muiri (Tillyard) 1926 (Permoscytina)
Permoscytina Tillyard 1926
kansasensis Tillyard 1926
Permopsylla Tillyard 1926
cubitalis (Carpenter) 1933 (Lithoscytina)
grandis Carpenter 1933
anomala Carpenter 1933
minuta Carpenter 1931
permiana Carpenter 1931
Paleoscytina Carpenter 1931
brevistigma Carpenter 1931
Glosselytrodea (Pterygota: Neoptera: Endopterygota) Perm.-Jur. (1 genus, 1 species)
Permoberothidae Tillyard 1932
Permoberotha Tillyard 1932 [= Dictyobiella Tillyard 1937]
villosa Tillyard 1932 [ = P. convergens Tillyard 1937; D. nervosa Tillyard, 1937]
Mecoptera (Pterygota: Neoptera: Endopterygota) Perm.-Hol. (6 genera, 8 species)
Permopanorpidae Tillyard 1926
Permopanorpa Tillyard 1926
formosa Tillyard 1926
inaequalis Tillyard 1926 [= P. tenuis, gracilis, dunbari, sellardsi, Tillyard 1926; P. raymondi Carpenter 1926]
schucherti Tillyard 1926
Permochoristidae Tillyard 1918 [= Mesochoristidae Tillyard 1926, and others]
Protochorista Tillyard 1926
tetraclada Tillyard 1926 [= P. pentaclada Tillyard 1926]
Protopanorpa Tillyard 1926
permiana Tillyard 1926
Agetopanorpidae Carpenter 1930
Agetopanorpa Carpenter 1930
maculata Carpenter 1930
Lithopanorpidae Carpenter 1930
Lithopanorpa Carpenter 1930
pusilla (Tillyard) 1926 (Protopanorpa)
Uncertain Family
Anormochorista Tillyard 1926
oligoclada Tillyard 1926
Uncertain Order (Pterygota: Neoptera) (8 genera, 9 species)
Uncertain Family
Apheloneura Carpenter 1976
minutissima Carpenter 1976
amplia Carpenter 1976
Choristosialis Tillyard 1932
enigmatica Tillyard 1932
Elmothone Carpenter 1976
martynovae Carpenter 1976
Gelasopteron Carpenter 1976
gracile Carpenter 1976
Nugioneura Tillyard 1937
problematica Tillyard 1937
Permembia Tillyard 1928
delicatula Tillyard 1928
Platychorista Tillyard 1926
venosa Tillyard 1926
Trachopteryx Carpenter 1976
martynovi Carpenter 1976
Summary:
Apterygota: 1 Order, 1 Family, 1 Genus, 1 Species
Palaeoptera: 6 Orders; 13 Families; 20 Genera; 39 Species
Neoptera: 10 Orders; 38 Families; 75 Genera; 113 Species
Total: 17 Orders; 52 Families; 96 Genera; 153 Species
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