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No. 9. The Amphiagrion sp. was identified by Huggins (1976) as distinct from A. saucium and A. abbreviatum, and therefore to be a new species. It remains to be described and named, and was not listed by Garrison. (The late Mrs. L.K. Gloyd studied Amphiagrion for many years. Rosser Garrison has told me that she had examined over 3000 specimens of the genus and was planning on describing the "mid-American" form (AZ to the Midwestern states) as A. mesonum. This name was in use as a manuscript name, and many workers during that time period therefore described Amphiagrion from the region as A. sp. Some workers today apparently consider Amphiagrion as a species with gradual clinal variation from east to west. The problem remains to be closed. Thanks to Rosser Garrison (Personal Communication, Sept. 1995) for discussing the issue with me.) In their recent master work on the Zygoptera of North America, Westfall and May (1996) state: "There is evidence that populations from the midwest, southwest into Arizona, probably repesent a third species, but their status is as yet unclear. Amphiagrion not clearly assignable to either A. abbreviatum or A. saucium are known from: AZ, CO, IL, IN, IA, KS, MI, NE, NM, ND, OK, SD, WI."No. 14. Argia immunda is added to the list based on a specimen from the USNM that was collected from Cherokee County on 27 April, 1963 by Gary F. Hevel (Male, determined by M. Davis, 1975)
No. 37. I have followed Garrison(1991) and Westfall & May (1996) by transferring Anomalagrion hastatum into the genus Ischnura as I. hastata (Say), 1839.
No. 41. Nehalennia gracilis is added to the list based on a photograph published in Insects in Kansas (2000, 3rd Edition, S.C. White & G.A. Salsbury, Kansas Department of Agriculture, Topeka, KS: p. 52). The photo was identified by D.G. Huggins and verified by Roy Beckemeyer. There is no locality record available for the photo other than it having been taken in the state (somewhere in the southeastern portion of Kansas).
No. 42. Telebasis salva is added to the list based on a specimen in the Florida State Collection of Arthropods that was the source for the listing by Westfall and May of this species in Kansas. Mike May and Bill Mauffray kindly tracked down the specimen, so the species is hereby added to the list vide Mauffray: A single envelope in the FSCA with the following data: "Telebasis salva Hagen 1 male, 1 female, Kansas, Sumner County, Hunters Mill Pond, 1 September 1936, Eldon Kile (Collector), Ex Williamson-Kennedy Collection, Ex M.J. Westfall Collection".
Update: Mark O'Brien of the Univ. of Michigan Museum of Zoology located 8 additional specimens that had been collected by Eldon Kile (same location) on Sept. 1, 1936, and 4 that were collected Sept. 11, 1936. All were determined by L.K. Gloyd. Mark has donated a pair of the specimens to us for the Kansas Biological Survey Collection.
Update #2: Betsy Betros of Lenexa, Kansas photographed T. salva (male) in Barber County, Kansas on 5 Sept. 1999. Photo id by R.J. Beckemeyer.No. 45. Aeshna interruptaq lineata is added to the list based on three male specimens in the Kansas University collection discovered by Ralph Charlton (1998). Labels carry the following information: Specimen #1: "T.F. Winburn coll / Manhattan, Ks. / July '29"; Specimen #2: "Coll. / W C Dick / Kansas / Marion Co. / Fall 1958"; Specimen #3: "Manh'tn, Ks. / '23 Sept. / Aeshna male / male app = lineata / color female = interna".
No. 65. Ophiogomphus rupinsulensis is known only from nymphal material for Kansas. Cook & Daigle (1985) determined that specimens previously identified as this species in Missouri and Arkansas were really a new species, O. westfalli. It is likely that the Kansas specimens are also of this new species but I have chosen to retain this entry as O. rupinsulensis until verification by examination of the specimens has been accomplished.
No. 74. Huggins listed and Garrison recognized both Macromia illinoiensis and M. georgina as valid species. However, T.W. Donnelly and K.J. Tennessen recently declared M. georgina to be a subspecies of M. illinoiensis, and the Kansas specimens they had inspected to be either intermediate forms (subspecies indeterminate) or of the subspecies M. i. georgina (1994, "Macromia illinoiensis and georgina: A Study of Their Variation and Apparent Subspecific Relationship (Odonata: Corduliidae)". Bull. of American Odonatology, 2(3):27-61). I have therefore removed Macromia georgina from the Kansas list.
No. 77-80. I follow Tennessen (1995, Personal Communication) who recommends Walker's treatment (1966, "On the generic status of Tetragoneuria and Epicordulia) Can. Entomol., 98:897-902); that is, making both Epicordulia and Tetragoneuria subgenera of Epitheca.
No. 77. Garrison, following the advice of K.J. Tennessen, synonomized Tetragoneuria williamsoni with T. costalis I have followed this advice (Tennessen, Personal Communication, 1995) as well, replacing the records of T. williamsoni with Epitheca costalis.
No. 79. While Paulson (1996, Personal Communication) and some other authorities consider Epitheca petechialis to be a synonym of Epitheca costalis, I follow the advice of Tennessen (1995, Personal Communication), and retain species status for E. petechialis pending further investigation.
No. 95. Erythrodiplax umbrata is added based on a specimen from the USNM collected by Gary F. Hevel on 11 July, 1964 from Oswego in Labette County (teneral male, determined by R.W. Garrison, 1986).
Update: A female specimen was collected in Sedgwick Co. by Roy J. Beckemeyer on 8 June, 1999.
In addition to the 57 species noted by the "^" annotation above, the following 7 species were also identified as occuring in Kansas by Kennedy (1917). However, Huggins (Huggins et al, 1976) was unable to substantiate the occurrence of the first 5 when he reviewed the material in the Kansas University collection. Because there are also no recent records of occurrence, we have listed them as Unsubstantiated:
Note that for Tramea
carolina (Linnaeus 1763). Kennedy referenced material from Pratt & Clark
Counties. Huggins determined that the Clark Co. specimen was T. onusta,
but could not locate the Pratt Co. specimen. Ragan Todd of Pittsburg, KS
(Crawford Co.) collected this species recently, thus
justifying adding it back onto the list.
In addition, Huggins (et al, 1976) determined the following specimens of
Kennedy to have been Misidentified:
Kennedy also listed a species which has subsequently been Synonomized:
Finally, the following species listed by Kennedy in 1917 are synonyms of
species in the current list:
Huggins (1975) reported that Allison's Crawford County collection had apparently been lost, and he did not comment further on Allison's listing of 34 species. Allison noted that "'The Insect Book', Howard; Hagen's 'Neuroptera'; 'The Dragon Flies of Indiana' by E.B. Williamson, were used in this work". There is no indication that he had any of his specimens verified by any contemporary authorities on Odonata. I list his 34 species and comment briefly on them here. Since there are no vouchers, the one species that would constitute and addition to the list is carried as Unsubstantiated.
1. Ęschna pentacantha [= Nasiaeschna pentacantha of this list
2. Ęschna verticalis [I strongly suspect this to be Aeshna constricta. Allison noted that the species was "Not rare", and A. constricta is present in Crawford County]
3. Anomalagrion hastatum [= Ischnura hastata (No. 37)]
4. Anax junius (No. 48)
5. Argia apicallis [sic] [= Argia apicalis] (No. 11)
6. Argia putridia [sic] [= Argia moesta (No. 15)]
7. Argia violacea [= Argia fumipennis violacea (No. 13)]
8. Celethemis [sic] elisa [= Celithemis elisa (No. 86)]
9. Celethemis [sic] eponina [=Celithemis eponina (No. 87)]
10. Dromogomphus spoliatus (No. 56)
11. Enallagma aspersum (No. 22)
12. Enallagma civile (No. 25)
13. Enallagma signatum (No. 30)
14. Epicordulia princeps [= Epitheca (Epicordulia) princeps (No. 79)]
15. Ischnura verticalis (No. 40)
16. Lestes unguiculatus (No. 8)
17. Libellua auripennis [There are no current collection records or known voucher species. None of the currently documented species of Libellula from southeast Kansas could be easily confused with auripennis. The species has been recorded in Oklahoma and Missouri, and could very well occur or have occurred in Kansas. I thus choose to add L. auripennis to the list of Unsubstantiated species of Odonata for Kansas.]
18. Libellula basalis [= Libellula luctuosa (No. 102)]
19. Libellula cyanea (No. 98)
20. Libellula pulchella (No. 104)
21. Libellula vibrans (No. 108)
22. Macromia taeniolata (No. 75)
23. Mesothemis simplicicollis [= Erythemis simplicicollis (No. 92)]
24. Pachydiplax longipennis (No. 110)
25. Pantala hymenaea (No. 112)
26. Pantala flavescens (No. 111)
27. Perithemis domitia [= Perithemis tenera (No. 113)
28. Plathemis lydia [=Libellula lydia (No. 103)]
29. Sympetrum albifrons [= Sympetrum ambiguum (No. 114)
30. Sympetrum corruptum (No. 115)
31. Sympetrum vicinum (No. 121)
32. Sympetrum semicinctum [Probably = Sympetrum occidentale fasciatum (No. 119), as all other Kansas specimens available that were labelled as semicinctum were the latter species.]
33. Tramea lacerata (No. 123)
34. Tramea onucta (No. 124)